The original plans for the toll road network in eastern OC did not include what is now CA 261. The tolled portion of CA 133 north of I-5 and what is now CA 241 between present CA 133 and CA 91 was designated as CA 231; CA 241 was where it is today southeast of the present 133/241 interchange. When the Irvine spur, originally intended to interchange with I-5 as well as empty out onto Jamboree Road into central Irvine, was first planned, the 231 designation was moved onto that facility and 241 was extended northeast from present 133 to the realigned 231. The original plans were to maintain the designations of the tolled facilities as completely separate entities from the existing non-tolled freeways -- which is why, in the original iteration, 133 wasn't simply extended north to 241 but changed at the I-5 junction to 231 (the intention was to keep all the tolled facilities northeast of I-5 as 2X1 numbers for system cohesion). But rather than sign the 133 extension as a short 2X1, it was decided to retain the 133 designation, figuring that segment to be more or less just an extra-long "feeder ramp" to 241. However, when it was decided to eliminate the 5/231 interchange near Jamboree in order to keep that tollway branch as a strictly Irvine server without the addition of I-5 traffic to the mix, signing the full section from Irvine to CA 91 as CA 231 was deemed inappropriate, as OCTA, the "parent" of the toll road authority, changed their priorities to emphasize the full CA 91 to I-5 at San Clemente routing as the "main trunk" (with the other segments being branches thereof). Now -- why OCTA and Caltrans didn't just maintain the Irvine "spur" as CA 231 is a matter of conjecture; Caltrans proposed "261" for the Irvine designation and OCTA "rubber-stamped" it, making the toll road "family" just as it is today. The last segments of the system to be done -- the 241 connector to CA 91 and the entirety of present 261 -- were opened to traffic in late 1998.
At that time (mid-late 1996) I was traveling monthly between Portland, OR and Anaheim Hills, CA; both my parents were ill at the time (my dad in early-stage Alzheimer's and my mom undergoing treatment for MDA*), so I was winding down my Portland time while working on my dissertation. The OCTA discussions concerning the configuration of the toll system were in full swing at the time; as some of the issues "dovetailed" with my dissertation topics, I decided to sit in on as many meetings, held at OCTA's Orange HQ, as I could while bringing my dad along just to keep his mind in action (after all, he was largely responsible for my being a roadgeek; I was simply returning the favor). Some of the discussions got a bit heated; in particular the one to eliminate the (eventual) 5/261 interchange; the rationale was that Irvine traffic heading toward Inland Empire housing was growing at such a high rate that a dedicated corridor eliminating much of the potential for intervening traffic from other arterial routes was considered optimal. The owner/developers of the shopping area at I-5 and Jamboree were up in arms about the interchange elimination; while they didn't reach their original goal of reinstating the tollway-to-freeway interchange, they did get major upgrades to the I-5/Jamboree interchange through the process (a partial victory for them), including expedition of traffic coming in both directions on the new CA 261 corridor to their businesses via changes in local ramp configurations. The final system configuration was presented in December 1996; CA 261 was opened to traffic 21 months later (which for CA must be some sort of record!). Of course, the whole system has never reached full fruition due to the truncation of the southern reaches of CA 241 for largely environmental reasons.
*MDA=myelodysplastic anemia; the bone marrow starts underproducing red blood cells. Still no cure; my mom got a few experimental treatments that, plainly, didn't work. Same disease that felled Carl Sagan.